Humulus japonicus (Japanese hops )

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1 of 8 9/24/2007 3:20 PM Home Early Detection IPANE Species Data & Maps Volunteers About the Project Related Information Catalog of Species Search Results :: Catalog of Species Search Humulus japonicus (Japanese hops ) Common Name(s) Full Scientific Name Family Name Common Family Scientific Name Images Synonyms Description Similar Species Reproductive/Dispersal Mechanisms Distribution History of Introduction in New England Habitats in New England Threats Early Warning Notes Management Links Documentation Needs Additional Information References Data Retrieval Maps of New England Plant Distribution COMMON NAME Japanese hops FULL SCIENTIFIC NAME Humulus japonicus Sieb. & Zucc FAMILY NAME COMMON Hemp family FAMILY SCIENTIFIC NAME Cannabaceae IMAGES Leaves and Inflorescences Fruits Close-up Habit Habitat Climbing Habit

2 of 8 9/24/2007 3:20 PM NOMENCLATURE/SYNONYMS Synonyms: Humulus scandens auct. non (Lour.) Merr. DESCRIPTION Botanical Glossary Humulus japonicus is a monoecious, annual vine that can grow between 0.5-2.5 m (1.5-8 ft.) in length. The stems and leaves of this plant have a very rough texture. The leaves are cordate in outline and palmately lobed. The leaves have 5 to 9 lobes separated by V-shaped sinuses. The margins of the lobes are serrulate and they have an acuminate apex. The leaves measure 5-12 cm (2-4.75 in.) in length, and the petiole is often longer than the leaf blade itself. The uppermost leaves are often 3-5 lobed. The staminate inflorescences are erect and measure 15-25 cm (6-10 in.) in length, while the pistilate inflorescences are cone-shaped spikes, 7-10 mm (0.25-0.4 in.) in size and the margins of the bracteoles are densely ciliate. The pistilate spikes are dull green in color and become abruptly acuminate. The flowers bloom in early to mid-summer. The achenes are yellow-brown, ovoid-orbicular in shape, and 4-5 mm (0.2 in.) in length. Page References Bailey 341, Fernald 556, Flora of North America 384, Gleason & Cronquist 73, Holmgren 70, Magee & Ahles 421. See reference section below for full citations. SIMILAR SPECIES Humulus lupulus L. (Common hops) Picture of H. lupulus Character Humulus japonicus Humulus lupulus Life cycle Annual Perennial Leaf lobes Apex of leaf lobes Petiole Bracteoles of pistilate spike 5-9 lobes Acuminate Longer than the blade Ciliate 3-7 lobed, sometimes unlobed Rounded Shorter than the blade Not ciliate REPRODUCTIVE/DISPERSAL MECHANISMS Humulus japonicus reproduces by seed. Its small seeds are dispersed by mechanical means, most notably by wind and by the moving water of rivers and streams.

3 of 8 9/24/2007 3:20 PM DISTRIBUTION Humulus japonicus is native to eastern Asia. In North America its range is from southern Quebec and Ontario to northern Georgia and west from North Dakota to Kansas. It is found in all of the states of New England. HISTORY OF INTRODUCTION IN NEW ENGLAND It is unclear when Humulus japonicus was first brought to North America, but it probably arrived here for intentional cultivation. In New England, the earliest records of its escape and naturalization come from eastern Massachusetts and southern Connecticut at the end of the nineteenth century. There are also records from several locations in Vermont (Brattelboro, Burlington, Bellows Falls) from the early part of the twentieth century. Most of these early records note that Humulus japonicus had escaped to "waste places." Robinson (1908) described Humulus japonicus as having "occasionally escaped from frequent cultivation." By 1950, Fernald characterized it as being found in waste places, roadsides and along fencerows from New England to Michigan and south from Virginia and Missouri. HABITATS IN NEW ENGLAND Abandoned Field Edge Open Disturbed Area River or Stream Roadside Vacant Lot Yard or Garden Humulus japonicus is often found in disturbed areas, along roadsides and utility right-of-ways, and in open fields, particularly when the soil is moist. It is also commonly found along river or stream banks and in floodplain areas, where it can form dense, almost continuous stands. THREATS When forming dense stands, Humulus japonicus can out compete native vegetation. DOCUMENTATION NEEDS Documentation required: A specific photograph or mounted snippet of the leaves with inflorescences. Best time for documentation: Summer, fall.

4 of 8 9/24/2007 3:20 PM ADDITIONAL INFORMATION

5 of 8 9/24/2007 3:20 PM Integrated Taxonomic Information System Taxonomic information about the species

6 of 8 9/24/2007 3:20 PM The PLANTS Database

7 of 8 9/24/2007 3:20 PM General information and map Flora of North America Extensive description and distribution map Delaware Wildflowers A close-up image of inflorescence REFERENCES Bailey, L.H. 1949. Manual of Cultivated Plants. Macmillan, New York. Britton, N.L. and A. Brown. 1970. An Illustrated Flora of the Northeastern United States vol. 1. Dover Publications Inc., New York. Fernald, M.L. 1950. Gray's Manual of Botany 8th ed. American Book Co., Boston. Flora of North America Editorial Committee, eds. 1993+. Flora of North America North of Mexico. Volume #3. Oxford University Press. Flora of North America Association ed. 2000. Flora of North America vol. 3. Oxford University Press, Oxford, New York. Gleason H.A. and A.C. Cronquist. 1991. Manual of Vascular Plants of the Northeastern United States and Adjacent Canada. 2nd ed. New York Botanical Garden, Bronx, New York. Holm, L.G., J.V. Pancho, J.P. Herberger, D.L. Plucknett. 1979. A Geographical Atlas of World Weeds. John Wiley and Sons, New York, USA. Holmgren N.H. 1998. Illustrated Companion to Gleason and Cronquist's Manual. New York Botanical Garden, Bronx, New York. Magee D.W and H.E. Ahles. 1999. Flora of the Northeast. University of Massachusetts Press, Amherst. USDA, NRCS. 2001. The PLANTS Database, Version 3.1. (http://plants.usda.gov). National Plant Data Center, Baton Rouge, LA 70874-4490 USA. DATA RETRIEVAL Select a task by clicking the radio button and then click "Subm Selection." Formatted display as table Export as comma-delimited text file Submit Selection MAPS OF PLANT DISTRIBUTION IN NEW ENGLAND

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