Plant Propagation Protocol for Eschscholzia californica ESRM 412 Native Plant Production

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Plant Propagation Protocol for Eschscholzia californica ESRM 412 Native Plant Production Source: Edward Step and William Watson Source: Henriette Kress

Map of North America Distrubution Source: USDA Plants Database Map of California Distribution Family Names Family Scientific Name: Family Common Name: Scientific Names Genus: Species: Species Authority: Variety: Sub-species: Cultivar: Map of Washington Distribution Source: USDA Plants Database TAXONOMY Papaveraceae Poppy Eschscholzia Eschlscholzia californica Cham. californica mexicana

Authority for Variety/Sub-species: Common Synonym(s) (include full scientific names (e.g., Elymus glaucus Buckley), including variety or subspecies information) Eschlscholzia californica var. crocea (Benth) Eschscholzia californica var. douglasii Eschscholzia californica var. maritime (E.L. Greene) Eschscholzia californica var. peninsularis (E.L. Greene) Eschscholzia californica procera Eschscholzia shastensis Common Name(s): Species Code (as per USDA Plants database): Geographical range (distribution maps for North America and Washington state) Ecological distribution (ecosystems it occurs in, etc): Climate and elevation range Local habitat and abundance; may include commonly associated species Plant strategy type / successional stage (stress-tolerator, competitor, weedy/colonizer, seral, late successional) Plant characteristics (life form (shrub, grass, forb), longevity, key characteristics, etc) California poppy, copa de ora or cup of gold, Mexican gold poppy. ESCA2 GENERAL INFORMATION Extends from southwestern Washington to Baja California, and south into western Texas, and as far as the coast. (See map above for distribution maps for North America and Washington State). Grassy and open areas, also grows in disturbed areas (such as along roadsides). Lower, warmer to full sun, often rocky elevations from sea level to no higher than about 2,000 ft.). Most of Cismontane CA, w. Mojave Desert & coastal OR. Also WA, NV, and N. MX; escaped freely elsewhere. Stress tolerator; tolerates dry, arid climates, likes low nutrient soils. Drought and full sun tolerant. Tolerates seaside conditions. Short-live perennials. Also found as herbs, annuals, and biennials. Herbaceous. Glabrous sometimes glaucous. Leaves are basal and occasionally cauline. Plants grow about 12-18 in. tall with a deep taproot. Tapered buds open into long-lasting flowers with 4 satin like petals reaching to about 2-3 in. in diameter each. Pollinated by beetles, but also taken over by European honey bee in other areas. After pollination, the leaves fall away leaving a ribbed seed pods, which then dry out, split, shooting out tiny black seeds. Cylindic fruit, dehiscent from base. PROPAGATION DETAILS Ecotype (this is meant primarily for Fort Funston, California (Young).

experimentally derived protocols, and is a description of where the seed that was tested came from): Propagation Goal (Options: Plants, Cuttings, Seeds, Bulbs, Somatic Embryos, and/or Other Propagules): Propagation Method (Options: Seed or Vegetative): Product Type (options: Container (plug), Bareroot (field grown), Plug + (container-field grown hybrids, and/or Propagules (seeds, cuttings, poles, etc.)) Stock Type: Time to Grow (from seeding until plants are ready to be outplanted): Target Specifications (size or characteristics of target plants to be produced): Propagule Collection (how, when, etc): Propagule Processing/Propagule Characteristics (including seed density (# per pound), seed longevity, etc): Pre-Planting Propagule Treatments (cleaning, dormancy treatments, etc): Growing Area Preparation / Annual Practices for Perennial Crops (growing media, type and size of containers, etc): Establishment Phase (from seeding to germination): Plants; Somatic Embryos Seed; gathering ripe pods just as they are beginning to split. Container (plug). (Young) Leach tube 15-30 days 12-18 inches tall. Root system: Firm plug in container. (Young). Collected during mid summer when pods are beginning to split open; placed in open container to dry out. Mature capsules are brown; seed is dark grey at maturity. (Young) Propagule 8 lbs. per acre at a depth of 1/8 to 1/4 inch. 250,000 seeds per pound. Seed cleaning: Shake seeds out of pods or rub pods to extract seeds. Place the pods in an open container to dry them out before removing/collecting the seeds. Bareroot grown, once seeds have been accumulated Seeds kept at room temperature. Seeds germinate without pre-treatment. Seeds placed in larger, open containers in a fully controlled greenhouse. (Young). Planted 6-8 inches apart. September or early fall Seeds are sown where plants are to stand, as they don t transplant well due to their deep taproot. Seeds are watered but not excessively; do relatively well on their own as they are significantly tolerant of disturbance climates. Do appreciate consistent moisture during germination.

Length of Establishment Phase: Active Growth Phase (from germination until plants are no longer actively growing): Length of Active Growth Phase: Hardening Phase (from end of active growth phase to end of growing season; primarily related to the development of cold-hardiness and preparation for winter): Length of Hardening Phase: Harvesting, Storage and Shipping (of seedlings): Length of Storage (of seedlings, between nursery and outplanting): Guidelines for Outplanting / Performance on Typical Sites (eg, percent survival, height or diameter growth, elapsed time before flowering): Other Comments (including collection restrictions or guidelines, if available): 28 days (Young) September to April Plants need to maintain sufficient moisture level, getting full sun. 31 weeks They often will withstand cold climates and will continue to bloom after the first frost. However, as autumn approaches, the plants should be cut from the ground and dried before the possibility of wilting approaches. 2-4 weeks Outplanting at an earlier age, for they do well on their own, re-seeding themselves. Outplanted into a well-drained soil. INFORMATION SOURCES References (full citations): See Below Other Sources Consulted (but that contained no pertinent information) (full citations): Protocol Author (First and last name): Kelsey Gaynor Middleton Date Protocol Created or Updated 4/24/11 (MM/DD/YY): References: Brach, A. R. & H. Song. 2006. efloras: New directions for online floras exemplified by the Flora of China Project. Taxon 55(1): 188-192. Calflora: Information on California plants for education, research and conservation, based on data contributed by dozens of public and private institutions and individuals, including the Consortium of Calif. Herbaria. [web application]. 2011. Berkeley, California: The Calflora Database [a non-profit organization]. Available: http://www.calflora.org/ (Accessed: May 30, 2011). California Native Plant Link Exchange: <http://www.cnplx.info/nplx/species?taxon=eschscholzia+californica>

efloras (2008). Published on the Internet http://www.efloras.org [accessed 22 February 2008]*' Missouri Botanical Garden, St. Louis, MO & Harvard University Herbaria, Cambridge, MA. Emery, Dara. Seed Propagation of Native California Plants. Santa Barbara, Botanic Garden. 1998. Lady Birdy Johnson. Wildflower Center. 2011. 4801 La Crosse Avenue, Austin, Texas, 78739. Linn, Willis, James C. Hickman. Jepson s Manual of the Flowering Plants of California. University of California: 1993. Munz P. A California Flora and Supplement. University of California Press, Berkeley and London, 1973. Pojar J., McKinnon A.1994 Plants of the Pacific Northwest: Washington, Oregon, British Columbia and Alaska, B.C. Ministry of Forests and Lone Publishing, Canada Young, Betty 2001. Propagation protocol for production of container Eschscholzia californica Cham. plants (Leach Tube);, San Francisco, California. In: Native Plant Network. URL: http://www.nativeplantnetwork.org (accessed 25 May 2011). Moscow (ID): University of Idaho, College of Natural Resources, Forest Research Nursery. See Appendix below.

Appendix: Original Protocol submitted by Maddie Scweitzer in 2005: Plant Data Sheet for Eschscholzia californica, California poppy Species Eschscholzia californica, California poppy Range USDA range 8 to 10 Climate, elevation 0-6561 ft. elevation in drier soils Local occurrence (where, how common)

Habitat preferences Full sun, grows excellent on slopes and roadsides Plant strategy type/successional stage (stress tolerator, competitor, weedy/colonizer, seral, late successional) Tolerates full sun, seaside conditions, and sand Associated species? May be collected as: (seed, layered, divisions, etc.) seed Collection restrictions or guidelines Flowers spring and throughout the summer and fall; allow seedheads to dry on plant and then remove and collect seeds Seed germination (needs dormancy breaking?) Seed dormancy generally absent in plants of cultivated heritage Seed life (can be stored, short shelf-life, long shelf life)? Recommended seed storage conditions? Propagation recommendations (plant seeds, vegetative parts, cuttings, etc.)

Sow seeds in fall or early spring directly into site as it is not easy to transplant Soil or medium requirements (inoculum necessary?) Well drained, drier soils; will tolerate poor soil; fertilizing is not necessary and over-fertilization will cause abundance of foliage that lacks flowers Installation form (form, potential for successful outcomes, cost) seed at approximately $1.59 per 2.5 grams, or $21.95 per pound Recommended planting density Space about 6-9 apart at a depth of 1/8 to 1/4 Care requirements after installed (water weekly, water once etc.) Water regularly to establish, and only an occasional deep watering is required thereafter; plants are drought-tolerant and over watering will cause the roots to rot Normal rate of growth or spread; lifespan One plant will grow up 12-20 inches and spread up to 20 inches; plant is stress deciduous and annual Sources cited http://www.laspilitas.com/plants/304.htm http://plants.usda.gov/cgi_bin/topics.cgi?earl=plant_profile.cgi&symbol =ESCA2 http://www.gardenguides.com/flowers/annuals/cpoppy.htm http://www.calflora.org/cgibin/species_query.cgi?special=calflora&where-calrecnum=3512&one=t Data compiled by Maddi Schweitzer on April 20, 2005